Davallia pentaphylla Blume, 1828

Description
Rhizome without the scales 2-4 mm in diametre, not white waxy (D. pentaphylla Rhizome). Scales nearly black with pale border quickly diminishing or disappearing towards the apex, distinctly acicular, often curling backward, bearing multiseptate hairs at least when young, peltate, 6-10 mm long. Stipes dark brown, adaxially grooved, 2-17 cm long, glabrous or with few scales. Lamina imparipinnate (D. pentaphylla Habitus3), leaflets entire or nearly so, occasionally lobed at the base or once branched (pairs of leaflets 2, occasionally 3-4), glabrous, not or slightly dimorphous. Leaflets entire or nearly so, sometimes with some basal lobes. Sterile terminal leaflet 2.5-16 cm long by 13-25 mm broad. Lateral leaflets 2.5-12 cm long by 8-25 mm broad. Margin flat or nearly so, distinctly crenulate to dentate at least towards apex. Petiolules 0-4 mm long. Leaflets of fertile leaves entire or nearly so, sometimes with some basal lobes. Fertile terminal leaflet 8-19 cm long by 7-15 mm broad. Lateral leaflets 4-14 cm long by 4-12 mm broad. Margins distinctly crenulate to dentate at least towards apex. Longest petiolules of fertile leaves 0-4 mm long. Pinnae narrowly ovate or linear (narrowly). Leaf axes glabrous. Margins of the lamina of each leaflet not thickened. Veins in sterile ultimate lobes parallel, once or twice branched from the base, reaching the margin. False veins not present. Sori separate (sometimes nearly connate, in a band along the margin) at the forking point of veins. Indusium also attached along the sides, pouch-shaped, oblong, longer than wide, 1.5-2.5 mm long by 0.75-2 mm broad, upper margin not elongated, truncate or slightly rounded, separated from to even with lamina margin (D. pentaphylla SEM, picture of indusia).

Distribution
Malesia: Sumatra (G. Kerinci, Bengkulu, Bangka, 5 coll.); Java (throughout many coll.); Lesser Sunda Islands (Bali many coll., Flores many coll., Sumbawa 2 coll.); Borneo (Sarawak, Lambir 1 coll., Kalimantan Selatan, G. Besar 1 coll., Kalimantan Timur, G. Beratus 1 coll., G. Medadam 1 coll.); Sulawesi (Northern 5 coll., Central 9 coll., Southwestern 6 coll.); Moluccas (Ternate 3 coll., Seram 6 coll.); New Guinea (many coll.).
Pacific: Bougainville (1 coll.); Solomon Islands (4 coll); Vanuatu (8 coll.); Fiji (many coll.).

Ecology
Epiphytic or epilithic on different kinds of rock, rarely terrestrial. Altitude 150-3200 m, but rarely on the lower altitudes.