Davallia corniculata

Davallia corniculata T. Moore, 1861

Description
Rhizome without the scales 3-4 mm in diametre, white waxy under the scales (D. corniculata Rhizome). Scales red-brown, without pale border, narrowed evenly towards the apex, often curling backward, not bearing multiseptate hairs, with marginal setae at least in distal part, peltate, 4-5 mm long by 0.5-1 mm broad. Stipes dark brown, adaxially grooved, 9-30 cm long, glabrous or with few scales. Lamina compound (D. corniculata Habitus), bipinnate or tripinnate towards base and in the middle part, deltoid and broadest towards base, glabrous, 16-50 cm long by 9-25 cm broad, not or slightly dimorphous. Longest petiolules 2-4 mm long. Pinnae narrowly ovate. Longest pinnae 5-19 cm long by 2-4.5 cm broad. Pinnules of at least the larger pinnae anadromous, linear oblong or narrowly ovate. Longest pinnules 12-25 mm long by 3-10 mm broad. Ultimate leaflets linear oblong, lobed almost to the midrib, or only shallowly lobed. Ultimate segments or lobes obtuse or acute without a tooth, or acute and usually ending in a tooth, 0.5-7 mm long by 1-2 mm broad. Upper ridge at the junction of the costa and pinna-rachis not swollen. Leaf axes glabrous. Margins of the lamina of each leaflet thickened and decurrent on the edge of the grooved rachis. Veins in sterile ultimate lobes pinnate, reaching the margin. False veins present. Sori separate, borne several on a segment, at the forking point of veins. Indusium attached at the base and only part of the sides, or also attached along the sides, pouch-shaped, more or less triangular to rhomboid or oblong, about as wide as long, c. 0.5 mm long and broad, upper margin not elongated, truncate or slightly rounded, separated from or even with lamina margin (D. corniculata SEM, picture of indusia). Lamina generally extending into a tooth at both sides or only at the outside of a sorus.

Distribution
Continental Asia: Southern Thailand (Nakawn Sritamarat, Khao Luang 3 coll.).
Malesia: Sumatra (Acèh, NW. of Takingeun 1 coll., Bengkulu, Seblat River 1 coll., East Coast, Bandar Baru 1 coll., N. Sibajak 1 coll., Tapanuli, Toba Plateau 1 coll.); Malay Peninsula (Pahang 2 coll., Penang 1 coll., Perak, Larut and Maxwell's Hill 5 coll.); Java (Western, G. Papandayan, G. Salak, Pasir Gombong, Telagawarna, Leuwiliang, Ciapus, Cibeurum, Cibodas, Cidadap, Cipanas, many coll.); Borneo (Sabah, Kinabalu, Dallas 1 coll.).

Ecology
Epiphytic or epilithic, sometimes in rather dry places. Altitude 300-1800 m.

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