Davallia wagneriana Copeland, 1905
Description
Rhizome without the scales 2-6 mm in diametre, white waxy under the scales (D. wagneriana Rhizome). Scales brown, red-brown, or nearly black, without pale border, narrowed evenly towards the apex, not or seldom curling backward, bearing (woolly) multiseptate hairs at least when young, peltate, 6-8 mm long by 1.5-2 mm broad. Stipes dark brown, adaxially grooved, 8-26 cm long, glabrous or with few scales. Lamina compound (D. wagneriana Habitus), bipinnate or pinnate with pinnatilobed to pinnatifid pinnae towards base and in the middle part, deltoid and broadest towards base or elongate, glabrous, 10-44 cm long by 5-20 cm broad, not or slightly dimorphous (but pinnulae of fertile leaves very narrow). Longest petiolules 1-4 mm long. Pinnae linear-triangular (curved upwards). Longest pinnae 4-13 cm long by 1.5-3 cm broad. Pinnules of at least the larger pinnae anadromous. Pinnules or pinnalobes linear oblong. Longest pinnules 10-15 mm long by 2-3 mm broad. Ultimate leaflets linear oblong, only shallowly lobed. Upper ridge at the junction of the costa and pinna-rachis not swollen. Leaf axes glabrous. Margins of the lamina of each leaflet not thickened. Veins in sterile ultimate lobes pinnate, reaching the margin. False veins present or not. Sori separate, borne several on a segment at the forking point of veins. Indusium also attached along the sides, pouch-shaped, oblong, longer than wide or about as wide as long, 1 mm long by 0.5-1 mm broad. Indusium upper margin not elongated, truncate or slightly rounded, separated from or even with lamina margin (D. wagneriana SEM, picture of indusia). Lamina generally extending into a tooth only at the outside of a sorus.
Distribution
Malesia: Sumatra (G. Kiermatuba 1 coll.); Borneo (Sarawak, G. Mulu 1 coll.; Kalimantan Timur 3 coll.); Philippines (Luzon, Panay 1 coll.; Mindanao 6 coll.; Negros 3 coll.; Leyte 1 coll.); Northern Sulawesi (4 coll.); Moluccas (Seram 4 coll.).
Ecology
Epiphyte in deep shadow (only scarcely recorded). Altitude 450-1600 m.